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The Turtles co-founder Mark Volman has died aged 78.
Guitarist and vocalist Volman passed away on September 5 after a 'brief, unexpected illness' in Nashville, Tennessee, his representative confirmed in a statement to People.
The Turtles were known for hits including Billboard number one single Happy Together in 1967 and Elenore in 1968.
Volman joined The Turtles - then known as the Nightriders and later the Crossfires - in 1963 when he was a student at Westchester High School in California with fellow founding member Howard Kaylan.
While Volman first started out as a saxophonist in the surf music band, he and Kaylan became lead vocalists when the group switched to folk rock.
The founding line-up of The Turtles became Volman, Kaylan, Don Murray, Al Nichol, Jim Tucker and Chuck Portz.
The band's later members were bassists Chip Douglas and Jim Pons, and drummers Joel Larson, Johnny Barbata, and John Seiter.
Their breakthrough single, a cover of Bob Dylan's It Ain't Me Babe was released in 1965 and reached the top 10 on the Billboard Hot 100.
This was followed by singles Let Me Be and You Baby. Tracks Grim Reaper of Love and Can I Get To Know You Better bombed in the US.
In 1966, after Murray and Portz quit, Barbata and Douglas entered the group.
Happy Together was released in 1967 and became the group's most famous single - kicking the Beatles' Penny Lane off the top of the US charts.
That same year Tucker quit the band.
In 1968 the band released a concept album called The Turtles Present the Battle of the Bands - which featured hit singles Elenore and You Showed Me.
Volman was known for his distinctive curly locks and that year, NME magazine even reported that he had insured his hair for $100,000.
Their final original album, Turtle Soup, was released in 1969 amid conflicts with the record label - who reportedly wanted to turn the band into a pop act like The Monkees.
After the band broke up in 1970, Volman and Kaylan joined Frank Zappa's band the Mothers of Invention, with Pons also becoming part of the line-up.
Due to being unable to use the Turtles' name due to a contract dispute with their label White Whale Records, Volman and Kaylan performed under the nicknames the Phlorescent Leech & Eddie or Flo & Eddie.
Volman and Kaylan played on several Zappa albums, and starred in his 1971 film 200 Motels.
They also sang backing vocals on T. Rex songs including 1971's Get It On (Bang a Gong) and the albums Electric Warrior and The Slider.
In 1992 Volman switched focus from music to study for a bachelor's degree at Loyola Marymount University, he graduated in 1997 aged 50.
Volman later earned a Master's degree in Fine Arts with an emphasis in screenwriting in 1999, also from Loyola.
He taught Music Business & Industry courses in the Communications and Fine Arts department at Loyola and also became an associate professor and coordinator of the Entertainment Industry Studies Program at Belmont University.
In 2015 Volman and Kaylan reunited as 'The Turtles ... Featuring Flo & Eddie' for their Happy Together Tour, marking the band's 50th anniversary.
Volman would be diagnosed with throat cancer that year and be declared cancer-free in 2016.
In 2020, Volman was diagnosed with Lewy body dementia (LBD), a progressive brain disorder that affects thinking, memory and movement.
He told People in 2023: 'I got hit by the knowledge that this was going to create a whole new part of my life. And I said, "OK, whatever’s going to happen will happen, but I’ll go as far as I can."'
'The challenges of this world affect everybody, and it’s been kind of fun being on the other side of a challenge like this and saying, "I feel good." My friends are here. I’m still here. And I want people to connect with me.'
Volman is survived by partner Emily Volman, his ex-wife Pat Volman, their daughters Hallie Volman and Sarina Miller and his brother Phil Volman.
His final Instagram post in January 2025 saw the frail star ask for strength for Los Angeles amid the devastating wildfires.
He wrote: 'As a born and raised Angeleno, I have watched all this in disbelief and heartbreak. One of the fires came dangerously close to my former Laurel Canyon home and friends have lost so much.
'Even if they didn’t lose their home, everyone in the city has lost a bit of faith and innocence. But, at my age & having been through a lot of shit, man, here’s what I know: No one can steal your spirit!
'We are of strong, formidable roots and even more creative, industrious minds. Los Angeles is Number One! I will never believe anything less.'